DR and FS mobilized the resources. All authors contributed to the research activities accom­plished in the project, including the fieldwork. JM, AS, GHD, DNO and FS collected from the study area, allotted collection numbers, pressed, and dried the voucher specimens. DNO and FS were assisted by GHD, JM and AS in species identification. DR performed the photos. GHD, DNO and KA analysed data and they wrote the man­uscript. KA assisted by DNO, DR, and FS super­vised the work.