MSS, LOS, TOR, AFMC, ACE and FHM collected and photographed the plant in the environment. MSS, RVCS and FHM wrote the text. MSS, TOR, FHM, RVCS and AMML revised the herbarium collections. MSS and RVCS identified the specimen and photographed it in the herbarium. AFMC, ACE, LOS, DOS and FHM revised the text.