LMN designed the study; BEA and JYU collected and preserved the specimen; BEA measured the specimen; LMN performed the genetic and morphological identification; ACA, AOO, AOA, ABO, EAE, SOO, OAU, ABA, AAU and CO provided technical assistance for the study and field collections; LMN, BEA, SOO and ACA wrote the initial draft of the manuscript, reviewed and finalized the manuscript; all authors read and approved the final manuscript.