New taxonomic and faunistic records of fungus gnats (Insecta, Diptera) from Montenegro, Romania, and Serbia

A total of 95 fungus gnats species were identified from sweep net samples collected from Montenegro, Romania and Serbia during 2010–2017. A mycetophilid species, Trichonta comis Gagné, 1981, is redescribed and a key is provided for the separation of European Boletina trivittata (Meigen, 1818)-group species. Ten species from Montenegro, 27 from Romania and 1 species from Serbia are reported for the first time. The presence of Stigmatomeria crassicornis (Stannius, 1831) in Romania is verified.


Introduction
The Sciaroidea is a species-rich superfamily, having 6 families occurring in the West Palaearctic region: Ditomyiidae, Diadocidiidae, Keroplatidae, Bolitophilidae, Mycetophilidae (often collectively treated as fungus gnats), Sciaridae (black-winged fungus gnats), and Cecidomyiidae (gall midges) (Ševčík et al. 2016). In this paper Sciaridae and Cecidomyiidae are not discussed. More than 1365 species have been described from the Palaearctic region (Bechev 2000) and new species are regularly found even from rather well-surveyed western Europe (Laštovka andŠevčík 2006, Kurina 2008). Larvae of European Sciaroidea are most often associated with microhabitats of humid forests, such as soil, dead wood, fruiting bodies of fungi and rarely plant tissues, eating mostly decomposing organic matter, while some genera are predaceous. However, some species are only met in wetlands or in subterranean habitats (Søli et al. 2000).
The first Romanian Sciaroidea records were published in the mid-19th century as a part of a larger monograph by Winnertz (1863). Later material collected from Romania was treated in the works of Strobl (1896), Thalhammer (1899), Lundström (1911aLundström ( , 1911bLundström ( , 1912Lundström ( , 1913Lundström ( , 1916, and Landrock (1914Landrock ( , 1925. Additions to the Romanian list of species were also provided by Tollet (1955), Decu-Burghele (1963), Burghele-Balacesco (1965, 1967, 1968, and Matile and Burghele-Balacesco (1969), based mostly on data about species found in caves. An additional publication added new information on the occurrence of Mycetophilidae in Romania (Sóos and Papp 1988). The expansion of knowledge on the Romanian fungus gnats fauna continued in the early 2000s during surveys of Carpathians and corridor of Danube (Pârvu 2005). This Carpathian survey was limited to the territory of the Maramureș Basin (Pârvu 2002(Pârvu , 2003 and Piatra Craiului National Park (Pârvu 2004a(Pârvu , 2004b. In the Checklist of Romanian Fauna (Moldovan et al. 2007), 205 species of fungus gnats (Ditomyiidae, Diadocidiidae, Keroplatidae, Bolitophilidae, Mycetophilidae) are listed though without any references and some previously reported species seem to have been missed out. The online resource Fauna Europaea (Chandler 2013) states that 206 species of Sciaroidea (except Sciaridae and Cecidomyiidae) are known from Romania. Both above mentioned checklists are deficient, not including all species reported previously from Romania. In the present paper we suggest that a total of 270 fungus gnats species are known from the country, previous to this paper and even this number should be regarded as an underestimate due to poor faunistic knowledge.
The Sciaroidea of Montenegro and Serbia has been much less investigated and no national checklist has hitherto been published. The last published checklist under the former Yugoslavia (focusing upon Serbia and Montenegro) by Bechev (1997) listed 56 fungus gnats species. Fauna Europaea (Chandler 2013) lists 166 sciaroid species (excluding Sciaridae and Cecidomyiidae), but this list combines the fauna of Montenegro and Serbia jointly. We suggest that a total of 168 fungus gnats species are known from these countries, previous to this paper.
The first author (L-PK) and his colleagues had collected insects from Montenegro, Romania, and Serbia during the years 2010-2017. Fungus gnats were sorted from the collected material and were sent to the second author (JS) for identification. Thus, the present paper aims to improve the faunistic knowledge of fungus gnats in the countries in question through providing an updated checklist of fungus gnats (Ditomyiidae, Diadocidiidae, Keroplatidae, Bolitophilidae, Mycetophilidae) recorded from Montenegro, Romania, and Serbia jointly.

Methods
The material was collected by sweep netting in Montenegro, Romania and Serbia (Fig. 1, Table 1), between 2010 and 2017, and identified by Jukka Salmela. All the material listed here, are stored in 96% ethanol and deposited in the natural history collection of the Regional Museum of Lapland, Rovaniemi, Finland (LMM). Individual catalogue numbers of the vouchers are given (e.g., DIPT-JS-2016-0301). The holotype of Trichonta comis Gagné, 1981 was loaned from Finnish Museum of Natural History (Zoological Museum) (MZHF), University of Helsinki, Helsinki, Finland. The arrangement of the treated families and genera follows Bechev (2000) and Fungus Gnats Online (http://sciaroidea.info/) that may better reflect the phylogeny of the families and subfamilies than Fauna Europea (Chandler 2013), which   classifies taxa according to alphabetical order.
In order to assess sample completeness of material collected from Romania, an individual rarefaction analysis was performed. Based on the analysis, the total number of studied specimens (131, x-axis) was plotted against the total number of observed species (86, y-axis). Data from Montenegro was omitted due to the rather small number of specimens collected and the remoteness of the sampling sites in the Carpathian Mountains. The analysis was calculated by using PAST 3.11 program (Hammer et al. 2001).
Morphological structures were examined using an Olympus CH2 microscope, Optika B-150 equipped with a Canon 650D camera and a LM Digital SLR Adapter. Layer photos were combined using the software Zerene Stacker.

Results
During this short and non-targeted survey 95 species collected from Montenegro, Romania, and Serbia (see Table 1) were identified. Our rarefaction curve based on material collected from Romania is still rising, meaning that the survey was incomplete (Fig. 2); more thorough faunistic study would yield more species from the area. Some rare and poorly known species were identified, such as Ditomyia fasciata (Meigen, 1818), Antlemon brevimanum (Loew, 1871), and Mycetophila curviseta Lundström, 1911 (Table 1).

Keroplatus reaumurii ssp. reaumurii
Macrorrhyncha rostrata (Zetterstedt, 1851) Material examined. A rather widespread European species, known from Central and Northern Europe and Russia (Chandler 2013), here reported for the first time from Romania. This species is, however, quite rare, being associated with pristine forests in Fennoscandia (Kjaerandsen andChandler 2011, Westling 2015); it is Red-Listed in Finland (Penttinen et al. 2010) and Sweden (Westling 2015). Immature stages are unknown, but most likely larvae of the genus dwell in decaying wood (Hutson et al. 1980). Bechev, 1989 Material examined. Monocentrota matilei was originally described from Bulgaria and Algeria (Bechev 1989). Papp (2003) reported this species from Hungary, and Chandler (2009) redescribed it based on specimens collected from Sardinia. The species is here reported for the first time from Romania. The species apparently has a fragmented range confined to mountainous areas of southeastern Europe and the Mediterranean area. Immature stages of M. matilei are unknown, but the European species M. lundstromi Edwards, 1925 is probably associated with decaying wood . Chandler, 1995 Material examined. The species was described from Slovakia (Chandler 1995) and has been later found from the Czech Republic (Ševčík 2004), Switzerland, Bulgaria (Bechev and Koç 2008), and Italy (Kjaerandsen et al. 2009) and is here reported for the first time from Romania. Immature stages are unknown, but larvae of Urytalpa probably dwell among soil, moss carpets, or decaying wood . Based on those few published localities, the species is most likely associated with European montane forests.

Family Bolitophilidae Winnertz, 1863
Bolitophila (Bolitophila) austriaca (Mayer, 1950) Material examined. A rather widespread Palaearctic species. In Europe B. austriaca is known from central and northern parts and Russia but is mostly absent or non-detected from a wide belt ranging from Greece to Lithuania (Chandler 2013). It is new for Romania and Montenegro. The species has been reared from a fruiting body of Tricholoma focale in Russian Karelia (Jakovlev 1994). (Curtis, 1836) Material examined. A widespread palaearctic species (Chandler 2013), here reported for the first time from Montenegro. It was reared from a number of agaric fruiting bodies (Jakovlev 1994), especially from Hypholoma fasciculare (Ševčík 2010). Winnertz, 1864 Material examined. It is a widespread Palaearctic species, known from several European countries (Chandler 2013), and new for the Romanian fauna. Larvae are polymycophagous, feeding on several agaric mushrooms (Jakovlev 1994(Jakovlev , 2011. According to Ševčík (2010), B. tenella is an uncommon species in the Czech Republic, perhaps preferring Pholiota species as host fungi. Polevoi, 1996 Material examined. It is a very rare and poorly known European species. The species was described from Russian Karelia (Polevoi 1996) and there are unpublished records from Norway and Sweden (Kjaerandsen 2012). Thus, B. melanoleuci is here reported for the first time outside Fennoscandia. Adults have been reared from the fruiting bodies of Melanoleuca melaleuca and M. brevipes (Polevoi 1996), and these fungi grow in grassy habitats, including man-made parks (Salo et al. 2016); the type material of the species was actually reared from mushrooms growing on city parks (Jakovlev 1993), as B. latipes Tollet, 1943). Edwards, 1913 Material examined. It is a widespread Palaearctic species, known from several European countries (Chandler 2013) and new to the Romanian fauna. It was reared from soft-bodied polypore fungi (Postia spp., Ševčík 2010). Records from agarics Hypholoma fasciculare and Pleurotus are reported (Jakovlev 2011) but may represent another Bolitophila species.

Bolitophila (Cliopisa) melanoleuci
Family Mycetophilidae Newman, 1834 Subfamily Mycomyinae Edwards, 1925 Mycomya (Mycomya) alpina Matile, 1972 Material examined.  Matile (1972) described this species from the French and Italian Alps. Later Väisänen (1984) redescribed this species and also reported it from Austria and Russian Far East (Kuril Islands). Thus, this species has a highly disjunct Palaearctic range. Mycomya alpina is here reported for the first time in Europe outside the Central European Alps. Immature stages are unknown, but Mycomya larvae are mostly associated with decaying wood and woodgrowing fungi (Väisänen 1984), and some species dwell in soil, litter, Carex-tussocks Roháček 2008, Jakovlev 2011) and even on (semi) aquatic lakeshores (Przhiboro 2012). (Dziedzicki, 1885) Material examined. This species has been reported from Central and northern Europe and Russia (Väisänen 1984, Chandler 2013) and is here reported as a new for the Romanian fauna. Immature stages are unknown. Väisänen, 1984 Material examined. This is a common and widespread Holarctic species but is new for the Romanian fauna. Larvae of the species are associated with fungal fruiting bodies (Väisänen 1984). (Zetterstedt, 1838) Material examined. A widespread Palaearctic species (Väisänen 1984), here reported for the first time from Montenegro. Larvae are mycophagous; adults have been reared from Leccinum scabrum (Väisänen 1984), Laxitextum bicolor, and Phlebia tremellosa (Jakovlev 2011).

Subfamily Sciophilinae Dziedzicki, 1885
Phthinia humilis Winnertz, 1864 Material examined. A rather widespread European species (Chandler 2013), here reported for the first time from Romania. The species is associated with forests; adults have been reared from decaying wood, moss carpet, and soil (Jakovlev 2011).
Subfamily Gnoristinae Edwards, 1925 Boletina joosti Plassmann, 1987 Material examined. This is a rare European species, hitherto known only from Germany. Plassmann (1986) described this species based on a holotype and a paratype males collected from Thuringia and Bavaria, respectively. The male specimens studied by Plassmann were collected in late April and late June. Instead, the specimen from Romania was collected in late August, and thus, B. joosti is on the wing during early and late season and adults probably overwinter. The closely related B. trivittata (Meigen, 1818) has a similar phenology pattern in Fennoscandia (J. Salmela, pers. obs.). Immature stages of B. joosti are unknown, but B. trivittata has been reared from rotting wood and soil litter; Boletina larvae are seldom found from fungal fruiting bodies (see Salmela et al. 2016 for a review). Boletina joosti is probably a very rare species, perhaps having a restricted and fragmented Central European range in mountainous areas.

Subfamily Leiinae Edwards, 1925
Docosia expectata Laštovka & Ševčík, 2006 Material examined. The species was described from the Czech Republic and Slovakia (Laštovka and Ševčík 2006), and is also known from Great Britain, Germany, Sweden and Finland (see Jakovlev et al. 2014). The species is new for Romanian fauna. Biology of D. expectata is still poorly known, but the species is perhaps associated with broadleaved forests (Kurina et al. 2004. The species is the most common and widespread Docosia in the Palaearctic region (Laštovka and Ševčík 2006), but not previously reported from Montenegro. Male hypopygium of D. gilvipes has been recently illustrated by Laštovka and Ševčík (2006) and Kurina (2008). Larvae are polymycophagous, known to occur in more than 40 species of fungi (Ševčík 2010).

Exechiopsis (Exechiopsis) unguiculata
(2011), citing Sasakawa and Ishizaki (1999). Immature stages of the species are unknown, but Exechiopsis larvae have been reared from both soil-dwelling agarics and saproxylic fungi (Jakovlev 2011). Exechiopsis unguiculata has been illustrated only by Lundström (1911a) and Zaitzev (2003), and thus photos of the male hypopygium are provided here. The male hypopygium is characterised by a ventroapical lobe of the gonocoxite that is narrowing apically, its apex rounded, margins straight and bordered by a black frame ( Figure 11). The medial appendage of the gonostylus is curved, long and narrow, bearing three subapical black spines arranged in a row and a long apical spine (Figs 12, 13). Dziedzicki, 1910 Material examined. The species has a rather restricted Central European range (Chandler 2013), here reported for the first time from Montenegro. An adult male of the species was collected from a cave in Romania (Burghele-Balacesco 1968, as R. matilei), but immature stages are unknown. Rymosia in general dwell in forests or wetlands (Chandler 1994), and larvae of some species are associated with fungal fruiting bodies (Jakovlev 1994). This species has been illustrated only twice (Dziedzicki 1910, Burghele-Balacesco 1968. Both illustrations are rather general, and photos of the male hypopygium of R. lundstroemi are provided here. Ventral lobe of gonostylus is short and black, truncated ( Figure 14); gonocoxites internally with prominent, black mesial hooks (Fig. 15); parameres about as long as cerci, having black, pointed apical spines (Figs 16, 17). (Stannius, 1931) Material examined. The species is probably very widespread in Europe (Chandler 2013) and has already been reported by Pârvu (2003Pârvu ( , 2004a from Romania. Stigmatomeria crassicornis is, however, close to S. obscura (Winnertz, 1864), a species that had been for decades considered as a junior synonym of S. crassicornis (see Kjaerandsen et al. 2007, Salmela andKaunisto 2015). Pârvu (2003) reported 2 females from Maramureș Depression, and the sex of the specimen examined from Piatra Craiului National Park was not specified (Pârvu 2004a). Because Pârvu's records may in principle also belong to S. obscura, the presence of S. crassicornis in Romania is verified herein. Larvae of the species may be associated with truffles (Edwards 1925, Chandler 2010), but Jakovlev (2011) reared the species from Ascocoryne sarcoides on birch log.  Winnertz, 1864 Material examined. Figure 1. Romania: Vârghiș, Vârghiș Canyon, hornbeam-beech forest, 630 m a.s.l., 46.2174° N, 025.5435° E, 5 May 2014, Török E. leg., 2 males, DIPT-JS-2016-0254. This is a rather widespread European species, here reported for the first time from Romania. Biology of the genus is poorly known, but S. gracilis has been reared from Mycoacia uda and Pleurotus dryinus (Chandler 1993), andZaitzev (2003) referred to a record from Coprinus fruiting bodies. Synplasta species are generally rare in the boreal zone, mainly associated with deciduous or old growth forests (J. Jakovlev, pers. comm.), and thus, S. gracilis is Red-Listed (NT) in Finland (Penttinen et al. 2010).

Synplasta gracilis
Synplasta rufilatera (Edwards, 1941) Material examined. This rare and poorly known European species was described from Great Britain (Edwards 1941, as Rhymosia) and was later recorded only from Poland (Zatwarnicki 2001, as Allodiopsis), Germany (Ševčík 2001), Finland (Jakovlev 2014), Sweden (Kjaerandsen et al. 2007), Russian Karelia, Ukraine and Siberia (Zaitzev 2003); it is here reported for the first time from Romania. Immature stages are unknown, but most likely it is a forest associated species. In Fennoscandia, this species is observed from both nemoral and north boreal vegetation zones (Kjaerandsen et al. 2007, J. Jakovlev pers. comm.). However, the species may consist of 2 taxa, proper S. rufilatera sensu Edwards and an undescribed taxon sensu Zaitzev (2003) (P.J. Chandler, pers. comm.). Chandler, 2001 Material examined. The species has a wide range, extending from Great Britain (Chandler 2000) and Portugal (Ribeiro 2003) to the Russian Far East (Zaitzev 2003). In Europe the species is also known from France (Chandler 2013), Finland (Jakovlev 2014), and Russian Karelia (Zaitzev 2003). This species is very close to M. gratiosa Winnertz, and some of the records published as M. gratiosa may actually belong to M. deflexa (e.g., Polevoi 2001) from eastern Finland). The species is Red-Listed in Finland (Penttinen et al. 2010), and it has been found from forests characterised by a high amount of dead woody material (J. Jakovlev, pers. comm.). Immature stages are unknown, but Mycetophila larvae live inside fungal fruiting bodies (Ševčík 2010(Ševčík , Jakovlev 2011. Caspers, 1984 Material examined. Figure 1 (Caspers 1984) and was later found in the British Isles (Chandler 1988), Bulgaria, Central Europe (Chandler 2013), Sweden, Norway, and Denmark (Kjaerandsen 2012). However, a recently described species, M. gemerensis Ševčík & Kurina 2011, is close to M. lastovkai, and thus older records of the latter species should be verified due to possible confusion of these 2 taxa (Salmela and Kaunisto 2015). Immature stages of M. lastovkai are unknown. Winnertz, 1864 Material examined. A widespread European species (Chandler 2013), here reported for the first time from Montenegro. Larvae are fungivorous (Zaitzev 2003;Ševčík 2010), most often reared from saproxylic fungi. (Dziedzicki, 1884) Material examined. A widespread European species (Chandler 2013), here reported for the first time from Romania. Mycetophila stylata has been reared once from a Lactarius fruiting body (Jakovlev 1994). Winnertz, 1864 Material examined. A widespread Holarctic species (Gagné 1975, Zaitzev 2003, here reported for the first time from Montenegro.
Legs. Coxae yellowish, apically infuscated, anterior edge of fore coxa bearing several light setae. Hind coxa with a posterobasal seta and hind tibia without posteroventral and anteroventral setae. Trochanters yellowish, ventrally black. Femora yellowish, with a narrow ventral band fading toward femoral apices. Tibiae and tarsi yellowish. Anteroapical depressed area of fore tibia ovate, bearing a comb of circa 26 hyaline setae. Length ratios of femur to tibia for fore-, mid-and hind legs: 1.0, 0.95, 0.88. Length ratios of tibia to basitarsus for fore-, midand hind legs: 1.24, 1.5, 1.62. Wing. Hyaline, length 3.2 mm. C and R veins dark brown, M1 and M2 mostly brown, CuA and M4 brown; veins r-m, first sector of M4, M1+2 and bases of M1 and M2 pale. These pale veins and Rs are also glabrous, bearing no stout, dark setae.
Abdomen. Brown, bearing light setae. Ninth tergite and cerci as in Figure 18. Gonocoxite dark brown, bearing numerous light setae (Fig. 19); ventrocaudal margin with a very modest median elevation. Gonostylus composed of two main lobes, dorsal and ventral (Fig. 20). Dorsal lobe (Fig. 21) is simple, lingulate, widest medially and narrowing apically; 14 stout setae on inner surface and four setae in a row on dorsal margin. Ventral lobe is intricate, consisting of the lobe itself and median projections. Apical half of the ventral lobe of gonostylus is rectangular; caudal outer corner is angular, bearing a dense dark setosity; inner corner with a finger-like projection and with a long subapical seta (Fig. 22); caudal margin even/truncated, with rather long pale setae. One of the projections of the ventral lobe is conspicuous, erect, dark brown; its stalk is widest basally, glabrous, narrowing toward apex; the apex is a beak-like structure, having a larger, pointed tooth and a comb consisting of ca 7-10 blunt teeth (Fig. 23). Aedeagus is rather wide, with a deep U-shaped median incision; apex truncated, rounded; caudal lateral corners with several sharp teeth (Fig. 24).
Female. Unknown. Biology. Immature stages of the species are unknown, but adults have often been collected from forests (see Taxon discussion below).
Taxonomic discussion. Holarctic species of the genus Trichonta were revised by Gagné (1981), and since then, 20 northern hemisphere species have been described (Chandler 1992, 2009, Bragina 1994, Chandler and Ribeiro 1995, Wu et al. 1995Zaitzev 2003, Hong et al. 2007. Trichonta comis is characterized by the presence of single posterobasal seta on hind coxa, absence of ventral setae on hind tibia, setosity of Cu petiole and hyaline wing lamina. In addition, the apical half of ventral lobe of gonostylus is bears a long seta. The aedeagus, not illustrated or verbally described by Gagné (1981), is also quite peculiar due to its spinose caudal corners. The type locality in Finland (Kuusamo, Juuma) is not known in detail, but is most likely within Oulanka National Park; the holotype male is the only specimen recorded from Finland. In Norway the species has been collected from both birch (Betula) (Søli 1994) and spruce (Picea) (Økland and Zaitzev 1997) dominated forests. The species is also known from the Czech Republic (Ševčík 1999, collected from a virgin beech forest) and Slovakia (Ševčík and Košel 2009b). It is new to the Romanian fauna. Trichonta comis is most likely a rare, boreo-montane forest-dwelling species, showing affinities to pristine forests. Larvae of Trichonta are predominantly associated with wood-decaying fungi (e.g., Jakovlev 2011) Trichonta vulcani (Dziedzicki, 1889) Material examined. Figure 1. Romania: Bălan, Hăşmaş Mt., Gálkút valley, spruce forest along a small brook, 1050 m a.s.l., 46.6493° N, 025.8415° E, 23 August 2014, Kolcsár L.-P. leg., 1 male, DIPT-JS-2016-0192.
A widespread Holarctic species (Gagné 1981, Zaitzev 2003, here reported as a new to the Romanian fauna. Immature stages are unknown.

Discussion
Based on individual rarefaction analysis, the obtained curve is still steadily rising, showing no signs of levelling off (Fig. 2). Thus, additional sampling and further examination of specimens is expected to increase the number of observed species. Hence, these results suggest a potentially species-rich, yet poorly researched, Sciaroidea fauna. In general, the lower Diptera fauna of Romania is still little investigated, evidenced by the discovery of several new species and new regional records (Ujvárosi and Krzeminska 2002, Ujvárosi et al. 2011a, 2011b, Török et al. 2013, Tkoč and Roháček 2014. Regarding the Carpathians, the systematic surveys performed so far suggest a diverse and poorly known Tipuloidea and Ptychopteridae fauna of this mountain range , and it is also definitely true in the case of Sciaroidea. Only 2 articles have been specifically dealt with the Carpathian Sciaroidea (Pârvu 2002(Pârvu , 2004a, and they added no less than 67 species to the Romanian fauna. All faunistic data (Table 1) and the actual checklist of fungus gnats (Ditomyiidae, Diadocidiidae, Keroplatidae, Bolitophilidae, Mycetophilidae) of Romania (Table 2) and Montenegro and Serbia jointly (Table 3) are available on TransDiptera Online Database (Kolcsár et al. 2017).
The Balkan area is a biodiversity hotspot, however, the Diptera fauna of the region is poorly known and has not been sufficiently explored (Ivković and Pont 2015). Only a few articles added new faunistic information to the native Diptera fauna of Montenegro and Serbia in the last 10 years (van Steenis et al. 2015, Płóciennik et al. 2014, Ivković et al. 2013, Stanković et al. 2014. No articles have been specifically dealt with the fungus gnat fauna of Montenegro or Serbia. Systematic, comprehensive and long-term surveys are necessary to expand our knowledge on the Montenegrin, Romanian, and Serbian dipteran fauna, including the highly diverse Sciaroidea. Long-term passive sampling devices like Malaise traps or bait traps should be used to explore phenology, population dynamics, and assemblage composition of different habitats. Increased faunistic knowledge will provide information about the formation and connection of the Montenegro, Romania, and Serbia fauna with surrounding areas.    (Edwards, 1925) Mycetophilidae Mycetophilinae Pseudexechia parallela (Edwards, 1925) Mycetophilidae Mycetophilinae Pseudexechia trisignata (Edwards, 1913) (Winnertz, 1864) Mycetophilidae Mycetophilinae *Synplasta rufilatera (Edwards, 1941) Mycetophilidae Mycetophilinae Tarnania dziedzickii (Edwards, 1941) Mycetophilidae Mycetophilinae Tarnania fenestralis (Meigen, 1818) Mycetophilidae Mycetophilinae Tarnania tarnanii (Dziedzicki, 1910) Bechev (1997) and Chandler (2013). New records for Montenegro are listed with one asterisk (*) and new for Serbia with two asterisks (**).