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Check List 21(5): 945-969
https://doi.org/10.15560/21.5.945 (16 Oct 2025)
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Check List 21(5): 945-969
doi: 10.15560/21.5.945
Received: 06 Nov 2024 | Approved: 20 Aug 2025 | Published: 16 Oct 2025
This article is part of:
Check List 21(5)
Authors
   
Alexander Griffin Bentley - Corresponding author
Waska Amazonía, Mera, Ecuador
Unidad de Investigación, Instituto Nacional de Biodiversidad, Quito, Ecuador
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Juan Pablo Reyes-Puig
Unidad de Investigación, Instituto Nacional de Biodiversidad, Quito, Ecuador
Fundación Red de Protección de Bosques ECOMINGA, Quito, Ecuador
Fundación Oscar Efrén Reye, Baños de Agua Santa, Ecuador
La Sapada Equipo Herpetológico, Finca La Argentina, Puyo, Ecuador
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Arley Benjamin Hidalgo
Waska Amazonía, Mera, Ecuador
Universidad Estatal Amazónica, Puyo, Ecuador
Universidad Regional Amazónica Ikiam, Tena, Ecuador
Fundación Cóndor Andino, Quito, Ecuador
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Josselin Alejandra Hernández Altamirano
Universidad Regional Amazónica Ikiam, Tena, Ecuador
Estación Biológica Pindo Mirador, Gobierno Autónomo Descentralizado Municipal de Pastaza, Mera, Ecuador
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María Elena Barragan-Paladines
Museo Fundación Herpetológica Gustavo Orcés, Quito, Ecuador
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Mario H. Yánez-Muñoz
Unidad de Investigación, Instituto Nacional de Biodiversidad, Quito, Ecuador
inabio.biodiversidad.gob.ec
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Diego Almeida-Reinoso
Escuela Politécnica Nacional, Quito, Ecuador
Centro de Conservación de Anfibios SAR-RANA, Llano Chico, Quito, Ecuador
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Mónica A. Guerra
Escuela Politécnica Nacional, Quito, Ecuador
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Jhony Arboleda
Universidad Técnica Particular de Loja, San Cayetano Alto, Ecuador
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Jaime Culebras
Photo Wildlife Tours, Quito, Ecuador
Fundación Cóndor Andino, Quito, Ecuador
https://www.photowildlifetours.com/
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Dione Alexandra Fiallos Moreno
Waska Amazonía, Mera, Ecuador
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Joaquín Sáenz
Waska Amazonía, Mera, Ecuador
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Philip M. Yoder
Rainforest Carpentry Inc., Sarasota, United States of America
Reserva Ecológica Alto Anzu, Mera, Ecuador
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Don Best
Waska Amazonía, Mera, Ecuador
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Ricardo Sebastián Vizcarra Vásconez
Waska Amazonía, Mera, Ecuador
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Manuel Genaro Peñafiel Flores
Town of Mera, Pastaza, Ecuador
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Alessandro Franchini
Southwestern Adventist University, Keene, United States of America
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Emma Carlson
Southwestern Adventist University, Keene, United States of America
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José Ignacio Segovia Larrea
Sapada Equipo Herpetológico, Finca La Argentina, Puyo, Ecuador
Universidad Estatal Amazónica, Puyo, Ecuador
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Ian Smitherman
Waska Amazonía, Mera, Ecuador
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Eric C.K. Gren
Southwestern Adventist University, Keene, United States of America
University of Montana, Hamilton, United States of America
Bitterroot Community Science Center, Hamilton, United States of America
Asclepius Snakebite Foundation, Centennial, United States of America
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Academic editor
Rafael de Fraga
Universidade Federal do Oeste do Pará, Santarém, Brazil
Instituto Nacional de Pesquisas da Amazônia, Brazil
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Author contributions
Conceptualization: AGB, JPRP, MEBP, DAFM, JA. Data curation: AGB, JAHA, DAR, MAG, EG. Formal analysis: AGB, IS. Funding acquisition: PMY. Investigation: AGB, JPRP, ABH, JAHA, DAR, MAG, MGPF, DAFM, JA, JS, PMY, DB, RSVV, AF, EC, JISL, EG. Methodology: AGB, JPRP. Resources: JAHA, MEBP, MYM, DAR, MAG, MGPF, DAFM, PMY. Supervision: AGB. Project administration: AGB. Validation: AGB, JPRP, MEBP, MYM, JC, EG. Visualization: AGB, DAFM, JPRP. Writing — original draft: AGB, ABH, JA, JS, PMY, DB, RSVV, AF, EC, JISL, EG. Writing — review and editing: AGB, JPRP, ABH, JAHA, MEBP, MYM, JC, IS, EG.
Conflict of interest
The authors have declared that no competing interests exist.
Supporting agencies
Waska Amazonía; Unidad de Investigación, Instituto Nacional de Biodiversidad; Fundación Red de Protección de Bosques ECOMINGA; Estación Biológica Pindo Mirador, Gobierno Autónomo Descentralizado Municipal de Pastaza; Museo Fundación Herpetológica Gustavo Orcés; Laboratorio de Herpetología, Departamento de Biología, Escuela Politécnica Nacional; Reserva Ecológica Alto Anzu; Manuel Genaro Peñafiel Flores (Town of Mera, Pastaza, Ecuador); Department of Biological Sciences, Southwestern Adventist University.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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