PN mobilized the resources. All authors contributed to the research activities accom­plished in the project, including the fieldwork. ACA, GHD, CGK and PN collected from the study area, allotted collection numbers, pressed, and dried the voucher specimens. ACA was assisted by GHD, CGK and PN in species identification. GHD and PN performed the photos. GHD and PN analysed data and wrote the man­uscript. ACA and PN super­vised the work.